List of pterosaur genera

This list of pterosaurs is a comprehensive listing of all genera that have ever been included in the order Pterosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (nomen dubium), or were not formally published (nomen nudum), as well as junior synonyms of more established names, and genera that are no longer considered pterosaurian. The list currently includes 288 genera.

Scope and terminology

There is no official, canonical list of pterosaur genera, but the most thorough attempts can be found at the Pterosauria section of Mikko Haaramo's Phylogeny Archive,^[1] the Genus Index at Mike Hanson's The Pterosauria,^[2] supplemented by the Pterosaur Species List,^[3] and in the fourth supplement of Donald F. Glut's Dinosaurs: The Encyclopedia series.^[4]

Authors and year

The authors column lists the authors of the formal description responsible for the erection of the genus listed. They are not necessarily the same as the authors of the type species as sometimes a species from one genus is determined sufficiently distinct to warrant the erection of a new genus to house it. If this is the case, only the latter authors will be listed. The year column notes the year the description was published.

Status

Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include:

Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type specimens are later assigned to the same genus, the first to be published (in chronological order) is the senior synonym, and all other instances are junior synonyms. Senior synonyms are generally used, except by special decision of the ICZN, but junior synonyms cannot be used again, even if deprecated. Junior synonymy is often subjective, unless the genera described were both based on the same type specimen.
Nomen nudum (Latin for "naked name"): A name that has appeared in print but has not yet been formally published by the standards of the ICZN. Nomina nuda (the plural form) are invalid, and are therefore not italicized as a proper generic name would be. If the name is later formally published, that name is no longer a nomen nudum and will be italicized on this list. Often, the formally published name will differ from any nomina nuda that describe the same specimen.
Preoccupied name: A name that is formally published, but which has already been used for another taxon. This second use is invalid (as are all subsequent uses) and the name must be replaced. As preoccupied names are not valid generic names, they will also go unitalicized on this list.
Nomen dubium (Latin for "dubious name"): A name describing a fossil with no unique diagnostic features. As this can be an extremely subjective and controversial designation, this term is not used on this list.

Age

The age column denotes the epoch of geologic time to which the fossils date. Genera that are invalid, misidentified, or otherwise do not represent a valid pterosaur are listed as age N/A because there was never a time in which a pterosaur by that generic name actually lived.

Location and notes

The location column designates the countries where remains of the relevant genus have been found. Genera that are invalid, misidentified, or otherwise do not represent a valid pterosaur are listed as location N/A because there was never a place in which a pterosaur by that generic name actually lived. The notes column is a collection of annotations on the scientific significance and taxonomic history of listed genera, as well as elaborations on the information presented in other columns.

The list

Genera

Genus	Authors	Year	Status	Age	Location	Notes
Aerodactylus^[5]	Vidovic & Martill	2014	Disputed	Late Jurassic	Germany	Some researchers suggest that every referred specimen is indistinguishable from Pterodactylus^[6]^[7]
Aerodraco^[8]	Holgado & Pêgas	2020	Valid	Early Cretaceous to Late Cretaceous	United Kingdom ( England)	Originally mistakenly thought to be a species of Pterodactylus
Aerotitan^[9]	Novas et al.	2012	Valid	Late Cretaceous	Argentina	The first azhdarchid known from South America
Aetodactylus^[10]	Myers	2010	Valid	Late Cretaceous	United States	Known from a partial lower jaw missing only two teeth
Afrotapejara^[11]	Martill et al.	2020	Valid	Late Cretaceous	Morocco	The first African tapejarid and the fourth pterosaur validly named from the Kem Kem Group
Aidachar^[12]	Nesov	1981	Misidentification	N/A	N/A	Actually a teleost fish whose remains were originally mistaken for jaw fragments of a ctenochasmatid pterosaur. The mistake was corrected five years later
Akharhynchus^[13]	Jacobs, Smith & Zouhri	2024	Valid	Early Cretaceous? to Late Cretaceous	Morocco	Based only on fragmentary premaxillae
Alamodactylus^[14]	Andres & Myers	2013	Valid	Late Cretaceous	United States	The holotype left wing is badly crushed
Alanqa^[15]	Ibrahim et al.	2010	Valid	Late Cretaceous	Morocco	Characterized by straight, pointed jaws, not unlike those of some azhdarchids
Albadraco^[16]	Solomon et al.	2020	Valid	Late Cretaceous	Romania	Has been suggested to represent a young Hatzegopteryx, but this was considered unlikely
Alcione^[17]	Longrich et al.	2018	Valid	Late Cretaceous	Morocco	Possessed proportionately short wings, which imply an adaptation either for a rapid flapping or to aid with underwater movement
Allkaruen	Codorniú et al.	2016	Valid	Early Jurassic	Argentina	Its braincase shows a number of features intermediate between earlier-diverging pterosaurs and more derived pterodactyloid taxa
Altmuehlopterus^[18]	Vidovic & Martill	2017	Valid	Late Jurassic	Germany	Formerly assigned to Germanodactylus
Amblydectes	Hooley	1914	Valid	Late Cretaceous	United Kingdom ( England)
Angustinaripterus^[19]	He et al.	1983	Valid	Middle Jurassic	China	May be piscivorous on account of its dentition
Anhanguera^[20]	Campos & Kellner	1985	Valid	Early Cretaceous to Late Cretaceous	Brazil Morocco	A 2003 study shows that it, alongside many other pterosaurs, held its head while facing the ground^[21]
Anurognathus^[22]	Döderlein	1923	Valid	Late Jurassic	Germany
Apatomerus^[23]	Williston	1903	Misidentification	N/A	N/A	Probably a misidentified plesiosaur^[2]
Apatorhamphus^[24]	McPhee et al.	2020	Valid	Late Cretaceous	Morocco	Possibly a chaoyangopterid although this cannot be confirmed due to the fragmentary nature of the remains
Aralazhdarcho^[25]	Averianov	2007	Valid	Late Cretaceous	Kazakhstan	May be the sister taxon to Phosphatodraco
Arambourgiania^[26]	Nessov	1989	Valid	Late Cretaceous	Jordan Morocco? United States?	Had very long cervical vertebrae, making its neck longer than that of a modern giraffe
Araripedactylus	Wellnhofer	1977	Dubious?	Early Cretaceous	Brazil
Araripesaurus	Price	1971	Dubious?	Early Cretaceous	Brazil
Archaeoistiodactylus	Lü & Fucha	2010	Valid	Middle Jurassic	China	Had quite short metacarpals
Arcticodactylus	Kellner	2015	Valid	Late Triassic	Greenland	Initially named as a species of Eudimorphodon, but it is distinct enough to be recognized as its own genus
Ardeadactylus^[27]	Bennett	2013	Valid	Late Jurassic	Germany	Possessed larger teeth than the contemporary Pterodactylus, suggesting it would have possibly preyed on larger fish
Argentinadraco	Kellner & Calvo	2017	Valid	Late Cretaceous	Argentina	Unusually, the top surface of the lower jaw consisted of ridges and depressions
Arthurdactylus	Frey & Martill	1994	Valid	Early Cretaceous	Brazil	Named after Arthur Conan Doyle, author of the 1912 novel The Lost World
Aurorazhdarcho^[28]	Frey, Meyer & Tischlinger	2011	Disputed	Late Jurassic	Germany	A 2025 analysis has suggested it may actually be a synonym of Gnathosaurus^[29]
Aussiedraco^[30]	Kellner, Rodrigues & Costa	2011	Valid	Early Cretaceous	Australia	Known solely from a fragmentary dentary
Austriadactylus^[19]	Dalla Vecchia et al.	2002	Valid	Late Triassic	Austria	Noted for a widened bony crest along the midline of the snout
Austriadraco	Kellner	2015	Valid	Late Triassic	Austria	Its uniquely fused frontals were once misidentified as a breast bone
Avgodectes^[31]	Peters	2004	Jr. synonym	N/A	N/A	Probable junior synonym of Haopterus
Aymberedactylus	Pêgas et al.	2016	Valid	Early Cretaceous	Brazil	The holotype jaw has a mixture of primitive and derived traits among tapejarids
Azhdarcho^[32]	Nesov	1984	Valid	Late Cretaceous	Kazakhstan Tajikistan? Uzbekistan	This genus is the namesake of the Azhdarchidae and Azhdarchoidea
Bakiribu	Pêgas et al.	2025	Dubious	Early Cretaceous	Brazil	Originally described as a ctenochasmatid pterosaur closely related to Pterodaustro,^[33] but may in fact be an amiid fish^[34]
Bakonydraco^[35]	Ősi, Weishampel & Jianu	2005	Valid	Late Cretaceous	Hungary	Has been variously suggested to be an azhdarchid, a tapejarid, or an azhdarchoid outside those groups
Balaenognathus^[36]	Martill et al.	2023	Valid	Late Jurassic	Germany	Was unable to fully close its mouth due to the upper jaw's strong curvature
Banguela^[37]	Headden & Campos	2015	Valid	Early Cretaceous	Brazil	Described as an unusually edentulous dsungaripterid, later suggested to be a species of Thalassodromeus,^[38] and finally recovered as a valid taxon, potentially a chaoyangopterid^[24]
Barbaridactylus^[17]	Longrich et al.	2018	Valid	Late Cretaceous	Morocco	Its femur has a slight distal expansion, a feature shared with Pteranodon
Barbosania^[39]	Elgin & Frey	2011	Valid	Early Cretaceous	Brazil	Some remains initially referred to Brasileodactylus may in fact belong to this genus
Batrachognathus^[40]	Ryabinin	1948	Valid	Late Jurassic	Kazakhstan
Beipiaopterus^[41]	Lü	2003	Valid	Early Cretaceous	China	Its wing membranes contained a large amount of blood vessels, suggesting they were possibly used in thermoregulation^[42]
Bellubrunnus^[43]	Hone et al.	2012	Valid	Late Jurassic	Germany	Uniquely for a pterosaur, it had forward-curving wingtips
Belonochasma^[2]	Broili	1939	Misidentification	N/A	N/A	Subsequently found to be a reptile of uncertain affinities
Bennettazhia	Nessov	1991	Valid	Early Cretaceous	United States	Originally assigned to the unrelated Pteranodon
Bergamodactylus^[44]	Kellner	2015	Jr. synonym	N/A	N/A	Junior synonym of Carniadactylus
Bogolubovia^[45]	Nessov & Yarkov	1989	Valid	Late Cretaceous	Russia	Has been suggested to be likely synonymous with Volgadraco, but it was considered valid based on newly discovered remains^[46]
Boreopterus^[47]	Lü & Ji	2005	Valid	Early Cretaceous	China	Probably used its needle-like teeth to trap small prey in a similar manner to Platanista river dolphins
Brachytrachelus	Giebel	1850	Preoccupied	N/A	N/A	Preoccupied name, now known as Scaphognathus
Brasileodactylus^[48]	Kellner	1984	Valid	Early Cretaceous	Brazil	One referred specimen indicates this genus had a thin crest in the lower jaw, contrasting the previous assumption that it had no crests in either jaw
Cacibupteryx^[49]	Gasparini, Fernández & de la Fuente	2004	Valid	Late Jurassic	Cuba	Named based on a three-dimensionally preserved skull
Caelestiventus^[50]	Britt et al.	2018	Valid	Late Triassic	United States	One of the largest pterosaurs known from the Triassic
Caiuajara^[51]	Manzig et al.	2014	Valid	Early Cretaceous	Brazil	Known from exceptionally large concentrations of fossils, implying a gregarious behavior for this taxon
Camposipterus^[52]	Rodrigues & Kellner	2013	Valid	Early Cretaceous	United Kingdom ( England)	The tip of the snout was flat
Campylognathoides^[19]	Strand	1928	Valid	Early Jurassic	Germany India?
Campylognathus^[53]	Plieninger	1894	Preoccupied	N/A	N/A	Preoccupied name, now known as Campylognathoides
Carniadactylus^[54]	Dalla Vecchia	2009	Valid	Late Triassic	Italy	Similar to Eudimorphodon, but differences in the size and dentition suggest distinct ecological roles for both taxa
Cascocauda^[55]	Yang et al.	2022	Valid	Middle Jurassic to Late Jurassic	China	Some of the preserved pycnofibres resemble the feathers of maniraptoran dinosaurs, which suggests a close relationship between these integuments
Cathayopterus^[56]^[57]	Wang & Zhou	2006	Valid	Early Cretaceous	China	The left side of the only described skull is damaged
Caulkicephalus^[58]	Steel et al.	2005	Valid	Early Cretaceous	United Kingdom ( England)	May have possessed a backward-facing head crest when alive
Caupedactylus^[59]	Kellner	2013	Valid	Early Cretaceous	Brazil	May have been relatively large for a tapejarid
Caviramus^[60]	Fröbisch & Fröbisch	2006	Valid	Late Triassic	Switzerland	Its discovery suggests a greater diversity of Triassic pterosaurs than previously thought
Cearadactylus^[61]	Leonardi & Borgomanero	1985	Valid?	Early Cretaceous	Brazil	Has been interpreted as a possible junior synonym of Brasileodactylus^[62]^[63]
Ceoptera^[64]	Martin-Silverstone et al.	2024	Valid	Middle Jurassic	United Kingdom ( Scotland)	May have inhabited a low-salinity lagoon environment
Changchengopterus^[65]	Lü	2009	Valid	Middle Jurassic	China	Two specimens are known, each belonging to different ontogenetic stages
Chaoyangopterus^[66]	Wang & Zhou	2003	Valid	Early Cretaceous	China	Originally mistakenly believed to be a nyctosaurid
Cimoliopterus^[52]	Rodrigues & Kellner	2013	Valid	Late Cretaceous	United Kingdom ( England) United States	Would have been capable of long-distance oceanic soaring due to its remains being mostly found in marine strata
Cimoliornis	Owen	1846	Valid	Late Cretaceous	United Kingdom ( England)
Coloborhynchus^[19]	Owen	1874	Valid	Early Cretaceous	United Kingdom ( England)
Comodactylus^[67]	Galton	1981	Dubious	Late Jurassic	United States	The first non-pterodactyloid pterosaur named from North America. Only a metacarpal is known
Cratonopterus^[68]	Jiang et al.	2023	Valid	Early Cretaceous	China	Exhibits traits unique among ctenochasmatids
Cretornis	Frič	1881	Valid	Late Cretaceous	Czech Republic
Criorhynchus	Owen	1874	Jr. synonym	N/A	N/A	Junior synonym of Ornithocheirus
Cryodrakon^[69]	Hone et al.	2019	Valid	Late Cretaceous	Canada	The northernmost pterosaur named from North America
Ctenochasma^[70]	Meyer	1851	Valid	Late Jurassic	France Germany
Cuspicephalus^[71]	Martill & Etches	2013	Valid	Late Jurassic	United Kingdom ( England)	The largest known wukongopterid and the only one currently described from outside China
Cycnorhamphus^[19]	Seeley	1870	Valid	Late Jurassic	France Germany
"Daitingopterus"^[72]	Maisch et al.	2004	Nomen nudum	N/A	N/A	Now referred to Altmuehlopterus
Daohugoupterus	Cheng et al.	2015	Valid	Middle Jurassic to Late Jurassic	China	The smallest pterosaur known from the Yanliao Biota
Darwinopterus^[73]	Lü et al.	2010	Valid	Middle Jurassic	China	Exhibits features of both basal pterosaurs and pterodactyloids
Dawndraco^[74]	Kellner	2010	Disputed	Late Cretaceous	United States	Possible junior synonym of either Geosternbergia or Pteranodon
Dearc^[75]	Jagielska et al.	2022	Valid	Middle Jurassic	United Kingdom ( Scotland)	Its discovery refutes the claim that pterosaurs did not reach larger body sizes until the appearance of the pterodactyloid forms of the Cretaceous
Dendrorhynchoides^[76]	Ji, Ji & Padian	1999	Valid	Early Cretaceous	China	Once thought to have come from the Jurassic. One of the youngest confirmed anurognathids
Dendrorhynchus^[77]	Ji & Ji	1998	Preoccupied	N/A	N/A	Preoccupied name, now known as Dendrorhynchoides
Dermodactylus	Marsh	1881	Dubious	Late Jurassic	United States
Dimorphodon^[19]	Owen	1859	Valid	Early Jurassic	United Kingdom ( England)
Diopecephalus	Seeley	1871	Valid	Late Jurassic	Germany
Dolicorhamphus^[78]	Seeley	1885	Valid	Middle Jurassic	United Kingdom ( England)
Domeykodactylus^[79]	Martill et al.	2000	Valid	Early Cretaceous	Chile	Combines features of both ctenochasmatids and dsungaripterids, but it was referred to as a member of the latter group
Doratorhynchus^[80]	Seeley	1875	Nomen vanum	Late Jurassic to Early Cretaceous	United Kingdom ( England)
Dorygnathus^[19]	Wagner	1860	Valid	Early Jurassic	France Germany
Douzhanopterus^[81]	Wang et al.	2017	Valid	Late Jurassic	China	Phylogenetically placed between the Wukongopteridae and Propterodactylus
Draigwenia^[82]	Holgado	2021	Valid	Early Cretaceous	United Kingdom ( England)	First described as a species of Ornithocheirus, but it is notably different from that taxon
Dsungaripterus^[19]	Young	1964	Valid	Early Cretaceous	China South Korea?
Elanodactylus^[83]	Andres & Ji	2008	Valid	Early Cretaceous	China	Its cervicals were convergently similar to those of azhdarchids
Eoazhdarcho^[84]	Lü & Ji	2005	Valid	Early Cretaceous	China	First described as an azhdarchid, though it has been found to be distantly related
Eopteranodon^[85]	Lü & Zhang	2005	Valid	Early Cretaceous	China	Potentially related to the coeval Sinopterus
Eosipterus^[19]	Ji & Ji	1997	Valid	Early Cretaceous	China	May have had bulky, elongated wings
Eotephradactylus^[86]	Kligman et al.	2025	Valid	Late Triassic	United States	The oldest pterosaur from North America
Epapatelo^[87]	Fernandes et al.	2022	Valid	Late Cretaceous	Angola	Known from incomplete, three-dimensionally preserved left limb bones belonging to two individuals
Eudimorphodon^[19]	Zambelli	1973	Valid	Late Triassic	Italy
Eurazhdarcho^[88]	Vremir et al.	2013	Valid	Late Cretaceous	Romania	Distinguished by some features of its cervical vertebrae
Eurolimnornis^[89]	Kessler & Jurcsák	1986	Dubious	Early Cretaceous	Romania	Originally described as a bird, but it was subsequently reinterpreted as a pterosaur^[90]
Europejara^[91]	Vullo et al.	2012	Valid	Early Cretaceous	Spain	The first tapejarid officially named from Europe
Faxinalipterus^[92]	Bonaparte et al.	2010	Misidentification	N/A	N/A	Although described as a pterosaur, it has since been reinterpreted as a non-pterosaurian archosaur, likely a lagerpetid^[93]
Feilongus^[94]	Wang et al.	2005	Valid	Early Cretaceous	China	Had a uniquely pronounced overbite. One of the largest basal pterodactyloids
Fenghuangopterus^[95]	Lü, Fucha & Chen	2010	Valid	Middle Jurassic	China	Only known from a fairly complete, although crushed, skeleton
Ferrodraco^[96]	Pentland et al.	2019	Valid	Late Cretaceous	Australia	Discovered by a cattle farmer while spraying herbicide
Forfexopterus	Jiang et al.	2016	Valid	Early Cretaceous	China	Unlike other ctenochasmatids, the tip of its snout was not spatula-shaped and possessed more curved teeth
Galgadraco^[97]	Giaretta et al.	2025	Valid	Late Cretaceous	Brazil	Related to Albadraco from Europe
Gallodactylus	Fabre	1974	Jr. synonym	N/A	N/A	Junior synonym of Cycnorhamphus
Garudapterus^[98]	Manitkoon et al.	2025	Valid	Early Cretaceous	Thailand	The first pterosaur named from Southeast Asia and the youngest confirmed gnathosaurine known to date
Gegepterus^[99]	Wang et al.	2007	Valid	Early Cretaceous	China	Had been speculated to be a juvenile of another ctenochasmatid
Geosternbergia	Miller	1978	Jr. synonym	N/A	N/A	Junior synonym of Pteranodon, specifically P. sternbergi
Germanodactylus^[19]	Yang	1964	Valid	Late Jurassic	Germany
Gladocephaloideus^[100]	Lü	2012	Valid	Early Cretaceous	China	The type specimen was originally believed to be an adult, but the discovery of the second fossil indicates it is actually much younger^[101]
Gnathosaurus^[19]	Meyer	1833	Valid	Late Jurassic	Germany United Kingdom ( England)
Gobiazhdarcho^[102]	Pêgas, Zhou & Kobayashi	2025	Valid	Late Cretaceous	Mongolia	One of the few pterosaurs described from Mongolia
Guidraco^[103]	Wang et al.	2012	Valid	Early Cretaceous	China	May have possessed a head crest with a slight front curve in life as suggested by the angled skull roof
Gwawinapterus	Arbour & Currie	2011	Misidentification	N/A	N/A	Initially thought to be a late-surviving istiodactylid pterosaur,^[104] but it was found to actually represent an indeterminate saurodontid fish the following year^[105]
Haliskia^[106]	Pentland et al.	2024	Valid	Early Cretaceous	Australia	The most completely known Australian pterosaur so far
Hamipterus^[107]	Wang et al.	2014	Valid	Early Cretaceous	China	Two different body and crest sizes are known, which may be a sexually dimorphic trait. Some well-preserved eggs are also known
Haopterus^[108]	Wang & Lü	2001	Valid	Early Cretaceous	China	The first Chinese pterosaur from which a skull is known
Harpactognathus^[109]	Carpenter et al.	2003	Valid	Late Jurassic	United States	Unusually for a rhamphorhynchid, it had a premaxillary crest on the midline of its skull that may have been expanded by soft tissue
Hatzegopteryx^[110]	Buffetaut et al.	2002	Valid	Late Cretaceous	Romania	Large and very robust in morphology
Herbstosaurus^[19]	Casamiquela	1975	Valid	Late Jurassic	Argentina
Hongshanopterus^[111]	Wang et al.	2008	Valid	Early Cretaceous	China	Initially placed within the Istiodactylidae by its describers, but it was later suggested to be in fact outside that family
Huanhepterus^[112]	Dong	1982	Valid	Early Cretaceous	China	The right half of its skeleton was destroyed by an explosion in a quarry
Huaxiadraco^[113]	Pêgas et al.	2023	Valid	Early Cretaceous	China	The third tapejarid known from the Jehol Biota. Originally referred to Huaxiapterus, which is now deemed synonymous with Sinopterus
Huaxiapterus^[114]	Lü & Yuan	2005	Jr. synonym	N/A	N/A	Junior synonym of Sinopterus
Iberodactylus^[115]	Holgado et al.	2019	Valid	Early Cretaceous	Spain	Possibly the largest pterosaur identified from the Iberian Peninsula
Ikrandraco^[116]	Wang et al.	2020	Valid	Early Cretaceous	China United Kingdom ( England)	Uniquely, its skull possessed a crest only in the lower jaw, hypothesized to have been an attachment point for a pelican-like gular pouch
Inabtanin^[117]	Rosenbach et al.	2024	Valid	Late Cretaceous	Jordan	The structure of its bones implies a capability of flapping its wings during flight in contrast to the soaring deduced for the much larger contemporary Arambourgiania
Infernodrakon^[118]	Thomas et al.	2025	Valid	Late Cretaceous	United States	Known from a single neck vertebra once believed to belong to Quetzalcoatlus
Ingridia^[119]	Unwin & Martill	2007	Jr. synonym	N/A	N/A	Objective junior synonym of Tupandactylus
Istiodactylus^[19]	Howse, Milner & Martill	2001	Valid	Early Cretaceous	China? United Kingdom ( England)	Had a short, broad skull with exceptionally large naso-antorbital fenestrae
Javelinadactylus	Campos	2021	Jr. synonym	N/A	N/A	Junior synonym of Wellnhopterus. Although named a few months earlier, its description was retracted over allegations that the describer did not have access to its holotype^[120]
Jeholopterus^[121]	Wang et al.	2002	Valid	Middle Jurassic to Late Jurassic	China North Korea?	Its large size potentially suggests it may have fed on fish other than insects as in other anurognathids
Jianchangnathus^[122]	Cheng et al.	2012	Valid	Middle Jurassic	China	Shared various features with Scaphognathus
Jianchangopterus^[123]	Lü & Bo	2011	Valid	Middle Jurassic	China	An almost complete skeleton is known, including the skull
Jidapterus^[124]	Dong, Sun & Wu	2003	Valid	Early Cretaceous	China	Possessed very small eye sockets
Kariridraco^[125]	Cerqueira et al.	2021	Valid	Early Cretaceous	Brazil	The type and only known skull was falsified by workers who glued it to the snout of an unrelated pterosaur
Kepodactylus^[126]	Harris & Carpenter	1996	Valid	Late Jurassic	United States	Discovered alongside the remains of Stegosaurus
Keresdrakon^[127]	Kellner et al.	2019	Valid	Early Cretaceous	Brazil	Found in direct association with the remains of Caiuajara, indicating the first known evidence for sympatry between pterosaurs
Klobiodon^[128]	O'Sullivan & Martill	2018	Valid	Middle Jurassic	United Kingdom ( England)	Its lower jaw had a gradual down curve at the front
Kryptodrakon^[129]	Andres, Clark & Xu	2014	Valid	Middle Jurassic to Late Jurassic	China	The oldest pterodactyloid currently named
Kunpengopterus^[130]	Wang et al.	2010	Valid	Middle Jurassic? to Late Jurassic	China	The species K. antipollicatus is noted for its opposable thumbs, an unusual trait among non-mammalian vertebrates^[131]
Lacusovagus^[132]	Witton	2008	Valid	Early Cretaceous	Brazil	The largest pterosaur of the Crato Formation. May be a chaoyangopterid
Laopteryx	Marsh	1881	Dubious	Late Jurassic	United States
Laueropterus^[133]	Hone	2026	Valid	Late Jurassic	Germany	Likely a close relative of the coeval Makrodactylus and Skiphosoura
Leptostomia^[134]	Smith et al.	2021	Valid	Early Cretaceous? to Late Cretaceous	Morocco	Potentially a probe-feeder as suggested by the unusual anatomy of its beak and the environment it would have lived in
Liaodactylus^[135]	Zhou et al.	2017	Valid	Late Jurassic	China	The oldest ctenochasmatid known to date. Had a shorter snout and fewer teeth than later members of that family
Liaoningopterus^[136]	Wang & Zhou	2003	Valid	Early Cretaceous	China	The fourth tooth of its upper jaw was the largest among pterosaurs
Liaoxipterus^[137]	Dong & Lü	2005	Valid	Early Cretaceous	China	Might have been an insectivore capable of protruding its tongue as indicated by the preserved hyoid
Limnornis^[138]	Kessler & Jurcsák	1984	Preoccupied	N/A	N/A	Preoccupied name, later renamed Palaeocursornis
Lingyuanopterus^[139]	Xu et al.	2022	Valid	Early Cretaceous	China	Some features of the skull are adaptations for a scavenging lifestyle
Linlongopterus^[140]	Rodrigues et al.	2015	Valid	Early Cretaceous	China	Its dentition looked quite similar to that of Haopterus and Mimodactylus
"Lithosteornis"^[141]	Gervais	1844	Nomen nudum	N/A	N/A	Referred to Ornithocheirus
Lonchodectes^[19]	Hooley	1914	Valid	Late Cretaceous	United Kingdom ( England)
Lonchodraco^[52]	Rodrigues & Kellner	2013	Valid	Early Cretaceous to Late Cretaceous	United Kingdom ( England)	Remains originally assigned to Pterodactylus and Lonchodectes
Lonchognathosaurus^[142]	Maisch, Matzke & Sun	2004	Valid	Early Cretaceous	China Russia?	Distinguishable by its pointed jaws
Longchengpterus^[143]	Wang et al.	2006	Valid	Early Cretaceous	China	The first istiodactylid known from China
Luchibang^[144]	Hone et al.	2020	Valid	Early Cretaceous	China	Has been found to be chimeric, with the tip of the snout being the actual holotype while the postcranial remains belong to an azhdarchomorph, probably Sinopterus^[145]
Ludodactylus^[146]	Frey et al.	2003	Valid	Early Cretaceous	Brazil	Had a long crest at the back of the skull, convergent with male Pteranodon longiceps
Luopterus^[147]	Hone	2020	Valid	Middle Jurassic to Late Jurassic	China	One of the smallest known pterosaurs. Originally believed to be a species of Dendrorhynchoides
Lusognathus^[148]	Fernandes et al.	2023	Valid	Late Jurassic	Portugal	Its teeth were robust and rounded or oval in cross-section
Maaradactylus^[149]	Bantim et al.	2014	Valid	Early Cretaceous	Brazil	Had one of the largest skulls of any anhanguerid
Macrotrachelus	Giebel	1852	Jr. synonym	N/A	N/A	Junior synonym of Pterodactylus
Makrodactylus^[150]	Hone et al.	2025	Valid	Late Jurassic	Germany	Smaller than most other known basal members of the Monofenestrata
Meilifeilong^[151]	Wang et al.	2023	Valid	Early Cretaceous	China	The most well-preserved and complete member of the Chaoyangopteridae yet known
Melkamter^[152]	Fernandes, Pol & Rauhut	2024	Valid	Early Jurassic	Argentina	The fifth pterosaur named from the Jurassic of South America
Mesadactylus^[153]	Jensen & Padian	1989	Valid	Late Jurassic	United States	Its sacral vertebrae unusually shared one trait with birds, which led to its misidentification as one
Microtuban^[154]	Elgin & Frey	2011	Valid	Late Cretaceous	Lebanon	Based on a juvenile specimen with many broken bones, which may have been caused by either a collision or an attack by a predator
Mimodactylus^[155]	Kellner et al.	2019	Valid	Late Cretaceous	Lebanon	Represents the first completely known Afro-Arabian pterosaur fossil. Had conical teeth at the front of its jaws
Mistralazhdarcho^[156]	Vullo et al.	2018	Valid	Late Cretaceous	France	Closely related to the South American Aerotitan
Moganopterus^[157]	Lü et al.	2012	Valid	Early Cretaceous	China	Known for having the longest skull of any toothed pterosaur
Montanazhdarcho^[158]	Padian, de Ricqlès & Horner	1995	Valid	Late Cretaceous	United States	Based on a mostly uncompressed fossil that may have belonged to a mature individual
Muzquizopteryx^[159]	Frey et al.	2006	Valid	Late Cretaceous	Mexico	The holotype fossil had been used as a decoration of an office wall
Mythunga^[160]	Molnar & Thulborn	2008	Dubious?	Early Cretaceous	Australia	Has been regarded as an indeterminate tropeognathine^[62]
Navajodactylus^[161]	Sullivan & Fowler	2011	Valid	Late Cretaceous	United States	Tentatively attributed to the Azhdarchidae by its authors, but the lack of pneumacity in its forelimbs does not support this
Nemicolopterus^[162]	Wang et al.	2008	Dubious	Early Cretaceous	China	Described as the smallest adult pterosaur yet recorded, but it has been recently suggested to represent an early development stage of Sinopterus^[113]
Nesodactylus^[163]	Colbert	1969	Valid	Late Jurassic	Cuba
Nesodon^[164]	Jensen & Ostrom	1977	Lapsus calami	N/A	N/A	Misspelling of Nesodactylus, also preoccupied by a toxodont
Nicorhynchus^[8]	Holgado & Pêgas	2020	Valid?	Early Cretaceous to Late Cretaceous	Morocco United Kingdom ( England)	A 2023 review claims both assigned species may be synonymous with Coloborhynchus, the genus to which they were initially attributed^[165]
Ningchengopterus^[166]	Lü	2009	Valid	Early Cretaceous	China	Preserves soft tissue remains, including wing membranes and pycnofibres
Nipponopterus^[167]	Zhou et al.	2025	Valid	Late Cretaceous	Japan	Similar in morphology to Gobiazhdarcho although it is also known from a subadult individual
Noripterus^[19]	Young	1973	Valid	Early Cretaceous	China Mongolia
Normannognathus^[142]	Buffetaut, LePage & LePage	1998	Valid	Late Jurassic	France	Has been recovered in a variety of phylogenetic positions within the Monofenestrata
Nurhachius	Wang et al.	2005	Valid	Early Cretaceous	China	Possibly paraphyletic as N. luei has been found to be closer to Hongshanopterus than to N. ignaciobritoi, the type species^[145]
Nyctodactylus^[168]	Marsh	1881	Jr. synonym	N/A	N/A	Junior synonym of Nyctosaurus
Nyctosaurus^[169]	Marsh	1876	Valid	Late Cretaceous	United States
Odontorhynchus	Stolley	1936	Jr. synonym	N/A	N/A	Junior synonym of Rhamphorhynchus
"Oolithorhynchus"^[2]	Whalley	2000	Nomen nudum	N/A	N/A	A manuscript name^[2] about which almost nothing is known
Ordosipterus^[170]	Ji	2020	Valid	Early Cretaceous	China	The morphology of its teeth may have made them not useful for cracking shells, unlike many other dsungaripterids
Orientognathus	Lü et al.	2015	Valid	Middle Jurassic	China	One of the basalmost rhamphorhynchids currently known
Ornithocephalus^[171]	Sömmering	1812	Jr. synonym	N/A	N/A	Junior synonym of Pterodactylus
Ornithocheirus^[19]	Seeley	1869	Valid	Early Cretaceous to Late Cretaceous?	Morocco? United Kingdom ( England)
Ornithodesmus^[2]	Seeley	1887	Misidentification	N/A	N/A	Later found to be a dromaeosaurid theropod dinosaur
Ornithopterus^[172]	Meyer	1838	Jr. synonym	N/A	N/A	Junior synonym of Rhamphorhynchus
Ornithostoma^[19]	Seeley	1871	Valid	Early Cretaceous	United Kingdom ( England)
"Osteornis"^[141]	Gervais	1844	Nomen nudum	N/A	N/A	Nomen ex dissertationae for pterosaur remains later named Palaeornis and Cimoliornis
Otogopterus^[173]	Ji & Zhang	2020	Valid	Early Cretaceous	China	Probably lived near braided rivers
Pachagnathus^[174]	Martínez et al.	2022	Valid	Late Triassic	Argentina	One of the only Triassic pterosaurs known from terrestrial environments
Pachyrhamphus^[172]	Fitzinger	1843	Preoccupied	N/A	N/A	Preoccupied name, now known as Scaphognathus
Palaeocursornis^[89]	Kessler & Jurcsák	1986	Dubious	Early Cretaceous	Romania	May be synonymous with Eurolimnornis
Palaeornis	Mantell	1844	Preoccupied	N/A	N/A	Preoccupied name, now called Rexarthuria
Pangupterus	Lü et al.	2016	Valid	Early Cretaceous	China	Its exact taxonomic placement is unknown
Paranurognathus^[175]	Peters	2005	Jr. snyonym	N/A	N/A	Junior synonym of Anurognathus
Parapsicephalus^[176]	Arthaber	1919	Valid	Early Jurassic	United Kingdom ( England)
Peteinosaurus^[19]	Wild	1978	Valid	Late Triassic	Italy
Petrodactyle^[177]	Hone et al.	2023	Valid	Late Jurassic	Germany	Large for a Jurassic pterosaur
Phobetor^[2]^[19]	Bakhurina	1986	Jr. snyonym	N/A	N/A	Junior synonym of Noripterus^[178]
Phosphatodraco^[179]	Pereda-Suberbiola et al.	2003	Valid	Late Cretaceous	Morocco	One of the few azhdarchid pterosaurs known from a mostly complete neck
Piksi^[180]	Varricchio	2002	Valid	Late Cretaceous	United States	Formerly erroneously described as a bird, although it was found to be a pterosaur by a later study^[90]
Plataleorhynchus^[181]	Howse & Milner	1995	Valid	Late Jurassic to Early Cretaceous	United Kingdom ( England)	Had a distinctive spoonbill-like beak with a rounded tip
Prejanopterus^[182]	Vidarte & Calvo	2010	Valid	Early Cretaceous	Spain	Had a bent snout contrasting with the straight lower jaw
Preondactylus^[19]	Wild	1984	Valid	Late Triassic	Italy	Was capable of developed flight despite having short wings
"Pricesaurus"^[183]	Martins-Neto	1986	Nomen nudum	N/A	N/A	A synonym of Anhanguera
Propterodactylus^[184]	Spindler	2024	Valid	Late Jurassic	Germany	Had been nicknamed the "Painten pro-pterodactyloid" before its formal description
Ptenodactylus^[185]	Seeley	1869	Preoccupied	N/A	N/A	Preoccupied name, referred to multiple pterosaur taxa
Ptenodracon^[186]	Lydekker	1888	Jr. synonym	N/A	N/A	Junior synonym of Pterodactylus
Pteranodon^[19]	Marsh	1876	Valid	Late Cretaceous	United States
Ptéro-dactyle	Cuvier	1809	Invalid	N/A	N/A	Renamed Pterodactylus
Pterodactylus	Cuvier	1809	Valid	Late Jurassic	Germany
Pterodaustro^[19]	Bonaparte	1970	Valid	Early Cretaceous	Argentina
Pterofiltrus^[187]	Jiang & Wang	2011	Valid	Early Cretaceous	China	Possessed a very elongated skull
Pteromonodactylus	Teryaev	1967	Jr. synonym	N/A	N/A	Junior synonym of Rhamphorhynchus
Pterorhynchus^[188]	Czerkas & Ji	2002	Valid	Middle Jurassic	China	Had both a low bony base and a large keratin extension forming its crest, a trait relatively unique among basal pterosaurs
Pterotherium^[189]	Fischer	1813	Jr. synonym	N/A	N/A	Junior synonym of Pterodactylus
Puntanipterus^[190]	Bonaparte & Sánchez	1975	Valid	Early Cretaceous	Argentina
Qinglongopterus^[191]	Lü et al.	2012	Valid	Middle Jurassic to Late Jurassic	China	Mostly similar to Rhamphorhynchus but with shorter wings and a smaller skull
Quetzalcoatlus^[19]	Lawson	1975	Valid	Late Cretaceous	United States
Radiodactylus^[14]	Andres & Myers	2013	Valid	Early Cretaceous	United States	May be the sister taxon to the Azhdarchidae^[17]
Raeticodactylus^[192]	Stecher	2008	Valid	Late Triassic	Switzerland	Had been suggested to be a synonym of the contemporary Caviramus^[54]
Rexarthuria^[193]	Thomas & McDavid	2026	Valid	Early Cretaceous	United Kingdom ( England)	Formally described after years of complicated taxonomy
Rhabdopelix^[2]	Cope	1870	Misidentification	N/A	N/A	At first thought to be a Triassic pterosaur, but it is now known to be, at least in part, a kuehneosaurid
Rhamphinion^[194]	Padian	1984	Valid	Early Jurassic	United States	Fragmentary fossils discovered in the United Kingdom initially attributed to the taxon have been found to actually come from a rhamphorhynchid
Rhamphocephalus	Seeley	1880	Misidentification	N/A	N/A	Later reassigned to the Thalattosuchia^[195]
Rhamphorhynchus^[19]	Meyer	1846	Valid	Late Jurassic	Germany Portugal? Spain? Tanzania? United Kingdom ( England)
Samrukia^[196]	Naish et al.	2012	Valid?	Late Cretaceous	Kazakhstan	Known only from a lower jaw once thought to belong to a bird. Possible synonym of Aralazhdarcho^[197]
Santanadactylus^[198]	De Buisonjé	1980	Dubious?	Early Cretaceous	Brazil	Would have been very large. Four species were named although they have been considered unrelated to each other
Saratovia^[199]	Averianov	2025	Valid	Late Cretaceous	Russia	One of the latest-surviving toothed pterosaurs
Scaphognathus^[19]	Wagner	1861	Valid	Late Jurassic	Germany
Seazzadactylus^[200]	Dalla Vecchia	2019	Valid	Late Triassic	Italy	Had an exclamation mark-shaped pteroid bone
Sericipterus	Andres et al.	2010	Valid	Late Jurassic	China	A close relative of Angustinaripterus
Serradraco^[201]	Rigal et al.	2018	Valid	Early Cretaceous	United Kingdom ( England)	Described as a possible lonchodectid, but a later analysis has suggested it was not a member of that family^[202]
Shenzhoupterus^[203]	Lü et al.	2008	Valid	Early Cretaceous	China	Can be distinguished by the arched head crest
Simurghia^[17]	Longrich et al.	2018	Valid	Late Cretaceous	Brazil? Morocco	Similar to the contemporary Alcione, although it is not represented by an adult
Sinomacrops^[204]	Wei et al.	2021	Valid	Middle Jurassic to Late Jurassic	China	Various bone tissue fragments have been lost as a result of their extremely fragile condition
Sinopterus^[205]	Wang & Zhou	2003	Valid	Early Cretaceous	China	A specimen reported in 2025 had gastroliths and phytoliths preserved in its stomach cavity, indicating an evidence for a herbivorous diet^[206]
Siroccopteryx^[207]	Mader & Kellner	1999	Valid?	Late Cretaceous	Morocco	Some researchers regard it as a probable junior synonym of Coloborhynchus
Skiphosoura^[208]	Hone et al.	2024	Valid	Late Jurassic	Germany	Transitional between basal pterosaurs and later pterodactyloids
Sordes^[19]	Sharov	1971	Valid	Late Jurassic	Kazakhstan
Spathagnathus^[209]	Fernandes et al.	2025	Valid	Late Jurassic	Germany	Unusually, its teeth were adapted for a diet of shelled prey items
Sultanuvaisia^[12]	Nessov	1981	Misidentification	N/A	N/A	Has been found to be actually a fish
Tacuadactylus^[210]	Soto et al.	2021	Valid	Late Jurassic	Uruguay	The holotype snout was formerly misidentified as belonging to a sawfish
Tapejara^[19]	Kellner	1989	Valid	Early Cretaceous	Brazil	Possessed a semicircular crest above its snout and a bony projection at the back of the head
Targaryendraco^[211]	Pêgas et al.	2019	Valid	Early Cretaceous	Germany	Originally named as a species of Ornithocheirus
Tendaguripterus^[212]	Unwin & Heinrich	1999	Valid	Late Jurassic	Tanzania	Inconsistent in phylogenetic placement
Tethydraco^[17]	Longrich et al.	2018	Valid	Late Cretaceous	Morocco	One study regarded it as an azhdarchid possibly synonymous with Phosphatodraco.^[213] However, this has been cast into doubt by subsequent research^[87]^[214]
Thalassodromeus^[215]	Kellner & Campos	2002	Valid	Early Cretaceous	Brazil	At first believed to have fed like extant skimmers, but it was found to have possessed strong jaw muscles, suggesting a ground foraging lifestyle
Thanatosdrakon^[216]	Ortiz David et al.	2022	Valid	Late Cretaceous	Argentina	May be the largest pterosaur of the Southern Hemisphere
Thapunngaka^[217]	Richards et al.	2021	Valid	Early Cretaceous	Australia	Its lower jaw is very similar to that of Anhanguera piscator
Titanopteryx^[218]	Arambourg	1959	Preoccupied	N/A	N/A	Preoccupied by a simuliid blackfly, later renamed Arambourgiania
Torukjara^[219]	Pêgas	2024	Valid	Early Cretaceous	Brazil	Remains originally identified as Caiuajara
Tribelesodon	Bassani	1886	Misidentification	N/A	N/A	Now known to be a misinterpreted specimen of the archosauromorph Tanystropheus
Tropeognathus	Wellnhofer	1987	Valid	Early Cretaceous	Brazil	One of the largest South American pterosaurs, with some azhdarchids rivaling it in size
Tsogtopteryx^[102]	Pêgas et al.	2025	Valid	Late Cretaceous	Mongolia	Small for an azhdarchid despite being fully grown
Tupandactylus^[220]	Kellner & Campos	2007	Valid	Early Cretaceous	Brazil	One specimen of T. imperator preserves pigment cells similar to those found in birds, as well as likely Stage IIIa feathers^[221]
Tupuxuara^[222]	Kellner & Campos	1988	Valid	Early Cretaceous	Brazil	Two valid species are known, eaching differing from each other in the crest morphology
Tylodorhynchus^[223]	Pêgas & Holgado	2026	Valid	Early Cretaceous	United Kingdom ( England)	Once named as a European species of Uktenadactylus
Uktenadactylus^[224]	Rodrigues & Kellner	2009	Valid	Early Cretaceous	United States	Formerly referred to Coloborhynchus
Unwindia^[225]	Martill	2011	Valid	Early Cretaceous	Brazil	Its snout was triangular in cross-section
Utahdactylus^[226]	Czerkas & Mickelson	2002	Valid	Late Jurassic	United States	Once considered indeterminate within the Diapsida^[227]
Vectidraco^[228]	Naish et al.	2013	Valid	Early Cretaceous	United Kingdom ( England)	Its ilium was distinctively pneumatized
Vesperopterylus^[229]	Lü et al.	2017	Valid	Early Cretaceous	China	Noted for having a reversed first toe, likely suggesting an arboreal lifestyle
Volgadraco^[230]	Averianov et al.	2008	Valid	Late Cretaceous	Russia	Some studies have proposed a nyctosaurid,^[17] pteranodontid,^[231]^[46] or basal pteranodontoid^[87] placement for this genus. However, a 2026 review restricted the taxon to the type snout and recovered it in the Azhdarchidae, supporting its original description^[232]
Wellnhopterus^[233]	Andres & Langston Jr.	2021	Valid	Late Cretaceous	United States	Would have hunted its prey in a raptorial manner as indicated by the blunt jaws and proportionally short neck
Wenupteryx^[234]	Codorniú & Gasparini	2013	Valid	Late Jurassic	Argentina	Possibly allied to the archaeopterodactyloids
Wightia^[235]	Martill et al.	2020	Valid	Early Cretaceous	United Kingdom ( England)	Known from damaged partial premaxillae
Wukongopterus	Wang et al.	2009	Valid	Middle Jurassic to Late Jurassic	China	The holotype may have broken its shin before its death
"Wyomingopteryx"^[236]	Bakker	1994	Nomen nudum	N/A	N/A	Likely intended to be coined for a pterosaur specimen with similarities to Istiodactylus
Xericeps^[237]	Martill et al.	2017	Valid	Late Cretaceous	Morocco	Possessed a unique forceps-like beak
Yelaphomte^[174]	Martínez et al.	2022	Valid	Late Triassic	Argentina	Had a tall, narrow bony crest similar to that of Raeticodactylus and Austriadactylus
Yixianopterus^[238]	Lü et al.	2006	Valid	Early Cretaceous	China	Known from a single, partially-preserved individual
Zhejiangopterus^[239]	Cai & Wei	1994	Valid	Late Cretaceous	China	Four specimens have been described so far
Zhenyuanopterus^[240]	Lü	2010	Valid	Early Cretaceous	China	Has been claimed to be the adult form of Boreopterus^[241]

Ichnogenera

Genus	Authors	Year	Status	Age	Location	Notes
Agadirichnus	Ambroggi & Lapparent	1954	Valid	Late Cretaceous	Morocco	The first pterosaur ichnogenus ever named
Haenamichnus^[242]	Hwang et al.	2002	Valid	Late Cretaceous	South Korea	Likely made by azhdarchids
Jinjuichnus^[243]	Jung et al.	2026	Valid	Early Cretaceous	South Korea	Preserved next to a trackway made by a small vertebrate, which indicates the trackmaker, probably a neoazhdarchian, was attempting to hunt the smaller animal
Pteraichnus	Stokes	1957	Valid	Late Jurassic	Asia^[244] Europe North America	Some tracks attributed to this ichnogenus have been considered crocodilian in origin by some scientists. However, recent studies have overturned those thoughts
Purbeckopus	Delair	1963	Dubious	Early Cretaceous	United Kingdom ( England)	May have been made by a relatively large pterosaur
Rhamphichnus^[245]	Mazin & Pouech	2020	Valid	Late Jurassic	Europe	Made by non-pterodactyloid pterosaurs

Oogenera

Although pterosaur eggs are known, some with complete embryos, no oogenera have been erected to house them. The holotype of the oospecies Oolithes sphaericus was briefly considered by Harry Govier Seeley to be pterosaurian in origin, although this attribution was dismissed before the formal erection of that oogenus.

References

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↑ Hanson, Mike (30 September 2006). "The Pterosaur Species List" (PDF). The Pterosauria. Archosauria.org. Archived from the original (PDF) on 29 July 2007. Retrieved 27 July 2007.
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↑ Kellner, A.W.A., and Campos, D.A. (1988). "Sobre un novo pterossauro com crista sagital da Bacia do Araripe, Cretaceo Inferior do Nordeste do Brasil. (Pterosauria, Tupuxuara, Cretaceo, Brasil)." Anais da Academia Brasileira de Ciências, 60: 459–469. [in Portuguese]
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↑ David M. Martill; Mick Green; Roy E. Smith; Megan L. Jacobs; John Winch (2020). "First tapejarid pterosaur from the Wessex Formation (Wealden Group: Lower Cretaceous, Barremian) of the United Kingdom". Cretaceous Research. 113 104487. Bibcode:2020CrRes.11304487M. doi:10.1016/j.cretres.2020.104487. S2CID 219099220.
↑ Bakker, R.T. (1994) Unearthing the Jurassic. In: Science Year 1995. World Book Inc.:Chicago, London, Sydney, Toronto, 76–89. ISBN 0-7166-0595-3.
↑ David M. Martill; David M. Unwin; Nizar Ibrahim; Nick Longrich (2017). "A new edentulous pterosaur from the Cretaceous Kem Kem beds of south eastern Morocco". Cretaceous Research. 84: 1–12. Bibcode:2018CrRes..84....1M. doi:10.1016/j.cretres.2017.09.006. hdl:2381/41058.
↑ Lü, J., Ji, S., Yuan, C., Gao, Y., Sun, Z. and Ji, Q. 2006. New pterodactyloid pterosaur from the Lowe Cretaceous Yixian Formation of Western Liaoning, p. 195–203. In: Lü, J., Kobayashi, Y., Huang, D. and Lee, Y. (eds), Papers form the 2005 Heyuan International Dinosaur Symposium. Beijing: Geological Publishing House.
↑ Cai, Z.; Wei, F. (1994). "On a new pterosaur (Zhejiangopterus linhaiensis gen. et sp. nov.) from Upper Cretaceous in Linhai, Zhejiang, China". Vertebrata PalAsiatica. 32: 181–194.
↑ Lü, J. (2010). "A new boreopterid pterodactyloid pterosaur from the Early Cretaceous Yixian Formation of Liaoning Province, northeastern Chian". Acta Geologica Sinica. 84 (2): 241–246. Bibcode:2010AcGlS..84..241L. doi:10.1111/j.1755-6724.2010.00204.x. S2CID 140600398.
↑ Witton, Mark P. (2013). "Boreopteridae". Pterosaurs: Natural History, Evolution, Anatomy. Princeton, NJ: Princeton University Press. pp. 164–169. ISBN 978-0-691-15061-1.
↑ Hwang, K.-G.; et al. (2002). "New pterosaur tracks (Pteraichnidae) from the Late Cretaceous Uhangri Formation, southwestern Korea". Geological Magazine. 139 (4): 421–435. Bibcode:2002GeoM..139..421H. doi:10.1017/s0016756802006647. S2CID 54996027.
↑ Jung, J.; Kim, K. S.; Xing, L.; Choi, B.-D. (2026). "New large pterosaur tracks from Korea and their implications on terrestrial behavior". Scientific Reports. 16 (1) 12363. Bibcode:2026NatSR..1612363J. doi:10.1038/s41598-026-48019-y. PMC 13087150. PMID 41991982.
↑ Kim, J. Y.; et al. (2006). "The oldest record of webbed bird and pterosaur tracks from South Korea (Cretaceous Haman Formation, Changseon and Sinsu Islands): More evidence of high avian diversity in East Asia" (PDF). Cretaceous Research. 27 (1): 56–69. Bibcode:2006CrRes..27...56K. doi:10.1016/j.cretres.2005.10.005.
↑ Mazin, Jean-Michel; Pouech, Joane (1 February 2020). "The first non-pterodactyloid pterosaurian trackways and the terrestrial ability of non-pterodactyloid pterosaurs". Geobios. 58: 39–53. Bibcode:2020Geobi..58...39M. doi:10.1016/j.geobios.2019.12.002.

External links

Pterosaur FAQs, by Raymond Thaddeus C. Ancog.
The Pterosaur Database, by Paul Pursglove.

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[3] Hanson, Mike (30 September 2006). "The Pterosaur Species List" (PDF). The Pterosauria. Archosauria.org. Archived from the original (PDF) on 29 July 2007. Retrieved 27 July 2007.

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[13] Jacobs, M. L.; Smith, R. E.; Zouhri, S. (2024). "A new ornithocheirid pterosaur (Pterosauria: Ornithocheiridae) from the mid-Cretaceous Ifezouane Formation, Kem Kem Group of Morocco". Cretaceous Research. 166 106015. 106015. doi:10.1016/j.cretres.2024.106015.

[LoneStarPterosaurs-14] 1 2 Andres, Brian; Myers, Timothy S. (2013). "Lone Star Pterosaurs". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 103 (3–4): 383–398. doi:10.1017/S1755691013000303. S2CID 84617119.

[ibrahimetal2010-15] Ibrahim, N.; Unwin, D.M.; Martill, D.M.; Baidder, L.; Zouhri, S. (2010). "A New Pterosaur (Pterodactyloidea: Azhdarchidae) from the Upper Cretaceous of Morocco". PLOS ONE. 5 (5) e10875. Bibcode:2010PLoSO...510875I. doi:10.1371/journal.pone.0010875. PMC 2877115. PMID 20520782.

[16] Alexandru A. Solomon; Vlad A. Codrea; Márton Venczel; Gerald Grellet-Tinner (2019). "A new species of large-sized pterosaur from the Maastrichtian of Transylvania (Romania)". Cretaceous Research. 110 104316. Bibcode:2020CrRes.11004316S. doi:10.1016/j.cretres.2019.104316. S2CID 213808137.

[newptero-17] 1 2 3 4 5 6 Nicholas R. Longrich; David M. Martill; Brian Andres (2018). "Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary". PLOS Biology. 16 (3) e2001663. doi:10.1371/journal.pbio.2001663. PMC 5849296. PMID 29534059.

[18] Steven U. Vidovic; David M. Martill (2017). "The taxonomy and phylogeny of Diopecephalus kochi (Wagner, 1837) and Germanodactylus rhamphastinus (Wagner, 1851)" (PDF). In D. W. E. Hone; M. P. Witton; D. M. Martill (eds.). New Perspectives on Pterosaur Palaeobiology (PDF). Vol. 455. The Geological Society of London. pp. 125–147. Bibcode:2018GSLSP.455..125V. doi:10.1144/SP455.12. S2CID 219204038. {{cite book}}: |journal= ignored (help)

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